1                    Ovenden P.M., Copland M.J.W. & Myers A.A. (1967)  Migrant Lepidoptera in Glamorganshire.  Entomologist's Record and Journal of Variation 79: 27

2                    King P.E., Richards J.G. and Copland M.J.W. (1968)  The structure of the chorion and its possible significance during oviposition in Nasonia vitripenis (Walker) (Hymenoptera: Pteromalidae) and other chalcids. Proceedings Royal Entomological Society London (A) 43: 13-2

3                    King P.E., Ratcliffe N.A. and Copland M.J.W. (1969)  The structure of the egg membranes in Apanteles glomeratus (L.) (Hymenenoptera: Braconidae).  Proceedings Royal Entomological Society London (A) 44: 137-142

4                    King P.E. and Copland M.J.W. (1969) The occurrence of Metoecus paradoxus L. (Col.: Rhipiphoridae) in Glamorgan.  Entomologist's monthly Magazine 105:114

5                    King P.E. and Copland M.J.W.  (1969) The structure of the female reproductive system in the Mymaridae (Chalcidoidea: Hymenoptera).  Journal of Natural History 3: 349-365

6                    King P.E., Copland M.J.W. and Hogge M.A F. (1970) Conservation begins at school. Fieldworker 1:25-28

7                    Copland M.J.W. and King P.E.  (1971) The structure and possible function of the female reproductive system in some Eulophidae and Tetracampidae. Entomologist 104: 4-28

8                    Copland M.J.W. and King, P.E.  (1971) The structure of the female reproductive system in the Chalcididae (Hym). Entomologist's monthly Magazine 107: 230-239

9                    Copland M.J.W. and King P.E.  (1972)  The structure of the female reproductive system in the Eurytomidae (Chalcidoidea: Hymenoptera) Journal of Zoology, London 166:185-212

10                Copland M.J.W. and King P.E.  (1972)  The structure of the female reproductive system in the Pteromalidae (Chalcidoidea:Hymenopterta) Entomologist 105: 77-96

11                Copland M.J.W. and King P.E.  (1972)  The structure of the female reproductive system in the Torymidae (Hymenoptera: Chalcidoidea) Transactions Royal entomological Society London 124: 191-212

12                Copland M.J.W., King P.E. and Hill D.S.  (1973)  The structure of the female reproductive system in the Agaonidae (Chalcidoidea: Hymenoptera). Journal Entomology (A) 48:25-35

13                Copland M.J.W.  (1976) Female reproductive system of the Aphelinidae (Hymenoptera: Chalcidoidea). International Journal Insect Morphology and Embryology 5:151-166

14                Copland M.J.W. and Askew R.R.  (1977) An analysis of the chalcidoid (Hymenoptera) fauna of a sand dune system. Ecological Entomology 2:27-46

15                Copland M.J.W.  (1979)  Hymenoptera in hop garden, with particular reference to parasitoids associated with the damson hop aphid Phorodon humuli. Entomologia experimentalis et applicata 25: 146-152

16                Galil J and Copland M.J.W.  (1981) Odontofroggatia galil Wiebes in Israel, a primary fig wasp of Ficus microcarpa L with a unique ovipositor mechansim (Epichysomallinae, Chalcidoidea). Proc Kon Ned Akad Wetensch (C) 884: 183-195

17                Ibrahim A.G. & Copland M.J.W. (1987)  Effects of temperature on the reproduction of Saisettia coffeae and its parasitoids.  Insect Science Applications  8: 351‑353

18                Avilla J & Copland M.J.W. (1987)  Effects of host stage on the development of the facultative autoparasitoid Encarsia tricolor (Hymenoptera: Aphelinidae).  Annals of Applied Biology 110: 381‑389

19                Avilla J & Copland M.J.W. (1988)  Development rate, number of mature oocytes at emergence and adult size of Encarsia tricolor at constant and variable temperatures.  Entomophaga 33: 289‑298

20                Tingle C.C.D. & Copland M.J.W.  (1988)  Effects of temperature and host plant on regulation of glasshouse mealybug (Hemiptera: Pseudococcidae) populations by introduced parasitoids (Hymenoptera: Encyrtidae).  Bulletin of Entomological Research 78: 135‑142

21                Tingle C.C.D. & Copland M.J.W.  (1988)  Predicting development of the mealybug parasitoids Anagyrus pseudococci, Leptomastix dactylopi and Leptomastidea abnormis under glasshouse conditions. Entomologia experimentalis et applicata 46: 19‑28

22                Tingle C.C.D. & Copland M.J.W.  (1989)  Progeny production and adult longevity of the mealybug parasitoids Anagyrus pseudococci, Leptomastix dactylopi and Leptomastidea abnormis (Hym.: Encyrtidae) in relation to temperature.  Entomophaga 34: 111‑120

23                Copland M.J.W. & Varley, M..J. (1990)  Glasshouse scale insects and biological control.  Orchid Review 98: 268‑270

24                Heidari M. & Copland M.J.W. (1992)  Host finding by Cryptolaemus montrouzieri (Col., Coccinelidae) a predator of mealybugs (Hom., Pseudococcidae).  Entomophaga 37: 621‑626

25                Cudjoe A.R., Neuenschwander P. & Copland M.J.W. (1992)  Experimental determination of the efficiency of indigenous and exotic natural enemies of the cassava mealybug, Phenacoccus manihoti Mat.‑ Ferr. (Hom., Pseudococcidae), in GhanaJournal of Applied Entomology 114: 77‑82

26                Heidari M. & Copland M.J.W. (1993)  Honeydew: A food resource or arrestant for the mealybug predator Cryptolaemus montrouzieriEntomophaga 38: 63‑68

27                Cudjoe A.R., Neuenschwander P. & Copland M.J.W. (1993) Interference by ants in biological control of the cassava mealybug Phenacoccus manihoti (Hemiptera: Pseudococcidae) in GhanaBulletin of Entomological Research 83: 15‑22

28                Croft P. & Copland M.J.W. (1994)  The influence of humidity on emergence in Dacnusa sibirica Telenga (Hymenoptera: Braconidae).  Biocontrol Science and Technology 4: 347‑351

29                Croft P. & Copland M.J.W. (1994)  Larval morphology and development of the parasitoid Dacnusa sibirica (Hym.: Braconidae) in the leafminer host Chromatomyia syngenesiaeEntomophaga 39: 85‑93

30                Croft P. & Copland M.J.W. (1995)  The effect of host instar on the size and sex ratio of the endoparasitoid Dacnusa sibericaEntomologia experimentalis et applicata 74: 121‑124

31                Healy T.P. & Copland M.J.W. (1995),  Activation of Anopheles gambiae mosquitoes by carbon dioxide and human breath.  Medical and Veterinary Entomology.  9: 331‑336

32                Ponsonby D. & Copland, M.J.W. (1995),  Olfactory responses by the scale insect predator Chilocoris nigritus. (F) (Coleoptera: Coccinellidae).  Biocontrol Science and Technology  5: 83‑93

33                Hemachandra K.S. & Copland M.J.W. (1995),  Aspects of reproductive biology of Delphastus pusillus, a coccinellid predator of glasshouse whitefly, Trialeurodes vaporariorum.  Tropical Agricultural Research 7: 48-55

34                Ponsonby D. & Copland, M.J.W. (1996),  Effect of temperature on development and immature survival in the scale insect predator, Chilocorus nigritus (F.) (Coleoptera: Coccinellidae).  Biocontrol Science and Technology 6: 101‑109

35                Islam K.S., Perera H.A.S. & Copland M.J.W. (1997), The effects of parasitism by an encyrtid parasitoid, Anagyrus pseudococci on the survival, reproduction and physiological changes of the mealybug, Planococcus citriEntomologia Experimentalis et Applicata 54: 77-83

36                Islam K.S. & Copland M.J.W. (1997),  Host preference and progeny sex ratio in a solitary koinobiont mealybug endoparasitoid, Anagyrus pseudococci (Girault), in response to its host.  Biocontrol Science and Technology 7: 449-456

37                Ponsonby D. & Copland, M.J.W. (1998),  Environmental influences on fecundity, egg viability and egg cannibalism in the scale insect predator, Chilocorus nigritus.  Biocontrol 43: 39-52

38                Gillani W.A. & Copland M.J.W. (1999), Defensive behaviour of the long-tailed mealybug Pseudococcus longispinus (Targioni Tozzetti) (Hemiptera: Pseudococcidae) against the brown lacewing Sympherobius fallax Navas (Neuroptera: Hemerobiidae) Entomologica Bari 33: 279-285

39                Islam K.S. & Copland M.J.W. (2000), Influence of egg load and oviposition time interval on the host discrimination and offspring survival of Anagyrus pseudococci (Hymenonoptera: Encyrtidae), a solitary endoparasitoid of citrus mealybug, Planococcus citri (Hemiptera: Pseudococcidae).  Bulletin of Entomological Research 90: 69-75

40                Karamaouna F. & Copland M.J.W. (2000),  Host suitability, quality and host size preference of Leptomastix epona and Pseudaphycus flavidulus, two endoparasitoids of the mealybug Pseudococcus viburni, and host size effect on parasitoid sex ratio and clutch size.  Entomologia Experimentalis et Applicata 96: 149-158

41                Ponsonby D.J. & Copland M.J.W. (2000), Environmental effects on the development and survival of the scale insect Abgrallaspis cyanophylli (Signoret) (Homoptera: Diaspidae) with reference to its suitability for use as a host for rearing biological control agents.  Biocontrol Science and Technology 10: 583-594

42                Healy T.P. & Copland M.J.W. (2000), Human sweat and 2-oxopentanoic acid elicit a landing response from Anopheles gambiaeMedical and Veterinary Entomology 14: 195-200

43                Sharpington P.J., Healy T.P. & Copland M.J.W. (2000),  A wind tunnel bioassay system for screening mosquito repellents.  Journal of the American Mosquito Control Association 16: 234-240

44                Ponsonby D.J. & Copland M.J.W. (2000), Maximum feeding potential of larvae and adults of the scale insect predator, Chilocorus nigritus with a new method of estimating food intake.  Biocontrol 45: 295-310

45                Karamaouna F. & Copland M.J.W. (2000), Oviposition behaviour, influence of experience on host size selection, and niche overlap of the solitary Leptomastix epona and the gregarious Pseudaphycus flavidulus, two endoparasitoids of the mealybug Pseudococcus viburni. Entomologia Experimentalis et Applicata 97: 301-308

46                Isikber A.A. & Copland M.J.W. (2001), Food consumption and utilisation by larvae of two coccinellid predators, Scymnus levaillanti and Cycloneda sanguinea, on cotton aphid, Aphis gossypiiBiocontrol 46: 455-467

47                Isikber A.A. & Copland M.J.W. (2002), Effects of various aphid foods on Cycloneda sanguinea. Entomologia Experimentalis et Applicata 102: 93-97

48                Healy T.P., Copland M.J.W., Cork A., Przyborowska A. & Halket J.M. (2002)  Landing responses of Anopheles gambiae elicited by oxocarboxylic acids.  Medical and Veterinary Entomology 16: 126-132


1                    Copland M.J.W.  (1973) Some mechanisms of sex determination in the parasitic Hymenoptera.  Proceedings of the Royal Entomological Society 38: 4 & 18-19

2                    Copland M.J.W.  (1983) Temperature constraints in the control of mealybug and scale insects. Bull IOBC/WPRS Darmstadt 1982: 142-145

3                    Ibrahim A.G. and Copland M.J.W. (1984), Biotic potential of Metaphycus helvolus. Proceedings 1984 British Crop Protection Conference 283-286

4                    Copland M.J.W.  (1986)  Role of temperature in biological control of tropical coccids in glasshouses. pp 221‑228 In Biological control in the tropics (Hussein M.Y.and Ibrahim A.G. eds) University Pertania Malaysia 516pp

5                    Copland M.J.W. & Varley M.J. (1987),  Progress in developing a controlled glasshouse environment to promote biological pest control.  IOBC Bulletin  10: 41‑45

6                    Croft P. & Copland M.J.W. (1993),  Size and fecundity of Dacnusa sibirica Telenga.  IOBC Bulletin 16: 53‑56

7                    Copland M.J.W., Perera H.A.S. & Heidari M. (1993),  Influence of host plants on the biocontrol of glasshouse mealybugs IOBC Bulletin 16: 44‑47

8                    Copland M.J.W. & Varly M.J. (2000),  Environments for Biocontrol Research.  Proceedings of UK Controlled Environment Users Group.  2000:17-21


1                    Copland M.J.W.  (1984) Scale insects on fruit MAFF Advisory leaflet No 88 8pp

2                    Copland M.J.W.  (1986) Earwigs friend or foe?  HDRA newsletter 22-23

3                    Varley M.J., Copland M.J.W., Wratten S.D. & Bowie M.H. (1994)  Parasites and Predators pp 33‑63 in Video techniqes in animal Ecology and Behaviour (Wratten, S.D. ed), Chapman & Hall, London, 211 pp

4                    Jervis M. & Copland M.J.W. (1996),  The Life Cycle  pp 63‑161  In Insect natural enemies: practical approaches to their study and evaluation.  Jervis, M. & Kydd, N. (Eds)  Chapman & Hall, London.  491 pp

5                    Ponsonby D. & Copland M.J.W. (1997),  Coccinellids and other Coleoptera. pp 29-60 In Soft scale insects their biology, natural enemies and control Vol 7B.  Ben‑Dov, Y. & Hodgson, C.J. (Eds) Elsevier, Amsterdam. 442pp


CABI (2001)  Crop Protection Compendium CD (Contributed section on Cryptolaemus montrouzieri)


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